Check the 5C socket contacts
The address and data lines reaching the EPROM socket at 5C are the one place a bad connection can hide after the buffers test good, because the EPROM sits in a socket. Inspect and if needed replace that socket.
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Source: Repairing DataEast 1987 (Laser War) to Sega 1995 (Batman Forever), Part Two, Clay Harrell (pinrepair.com)