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Which CPU boards can run which system firmware

A System 3 board can be modified to run System 3, 4 and most System 6 code (not Firepower). System 4 boards run System 3, 4 and most System 6 code, and System 6 boards run System 3 to 6 code; none of these run System 7 code. A System 7 board runs anything from System 3 to 7 when its ROM jumpers are set and the EPROMs are formatted to match.

Systems

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Source: Repairing Williams System 3 to System 7 Pinball 1977 to 1984, Part One, Clay Harrell (pinrepair.com)